At one point, Firefox (3.5 specifically) was #1, for a brief moment:
> Between mid-December 2009 and February 2010, Firefox 3.5 was the most popular browser (when counting individual browser versions) according to StatCounter, and as of February 2010 was one of the top 3 browser versions according to Net Applications. Both milestones involved passing Internet Explorer 7, which previously held the No. 1 and No. 3 spots in popularity according to StatCounter and Net Applications, respectively - https://en.wikipedia.org/wiki/Firefox_3.5
Then Chrome appeared and flattened both IE and Firefox.
> Between mid-December 2009 and February 2010, Firefox 3.5 was the most popular browser (when counting individual browser versions) according to StatCounter, and as of February 2010 was one of the top 3 browser versions according to Net Applications. Both milestones involved passing Internet Explorer 7, which previously held the No. 1 and No. 3 spots in popularity according to StatCounter and Net Applications, respectively - https://en.wikipedia.org/wiki/Firefox_3.5
Then Chrome appeared and flattened both IE and Firefox.